Rapunzel, having asked Mother Gothel on the day before her eighteenth birthday if she could leave the tower and see the floating lights that appeared annually, soon realizes she will never be freed and escapes with the help of a thief named Flynn.
Separated from his own species as a hatchling and raised by a clan of lemurs, Aladar's life is plunged into chaos when a meteor shower forces him to join a group of migrating dinosaurs searching for a safe nesting ground.